As a test of the FDR procedure, I will revisit the case of N=9, M=9. (It now occurred to me that I was not careful about checking for repeated edges in the underlying directed edge construction. Furthermore, it is possible to get n_i \to n_j and n_j \to n_i, in which case, the "true" number of inferrable undirected edges will be less than the nominal M.)
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